Gold Skull Locations:
Black Eye (Located: Crow's Nest)
When activated, you will have to battle enemies with melee tactics to regain shield energy.
Catch (Located: The Storm)
When activated, enemies will throw more grenades.
Famine (Located: The Ark)
When activated, dropped weapons will hold less ammunition.
Fog (Located: Floodgate)
When activated, the motion tracker disappears.
Iron (Located: Sierra 117)
When activated, you will be sent back to the beginning of the stage you're on when you die instead of to the last checkpoint.
Mythic (Located: Halo)
When activated, all enemies will have twice as much health.
Thunderstorm (Located: The Covenant)
When activated, enemies will be of a higher rank.
Tilt (Located: Cortana)
When activated, enemies will become more resilient to your attacks.
Tough Luck (Located: Tsavo Highway)
When activated, enemies are skilled at getting away from projectiles
such as grenades.
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Silver Skull Locations:
Blind (Located: Sierra 117)
The arms of the player and the motion tracker disappear.
Cowbell (Located: The Ark)
Your explosives increase in strength.
Grunt Birthday Party (Located: Crow's Nest)
If you manage to get a headshot on a Grunt, the Grunt will explode.
I Would Have Been Your Daddy (Located: The Covenant)
Unlocks new dialogue for the campaign.
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